"Parse error: syntax error, unexpected T_ENDWHILE"

I was updating some pages on my website, and when I tried to see if all was well, I got this error message:

Parse error: syntax error, unexpected T_ENDWHILE in /home/…/page.php on line 22
This is page.php

<?php get_header(); ?>

<p align="center">
	<span class="title">
		<?php if (have_posts()) : while (have_posts()) : the_post(); ?>
		<id="post-<?php the_ID(); ?>"><?php the_title(); ?>
	</span>

	<div class="news">
		<?php the_content('Leia o restante da página &raquo;'); ?>
		<?php link_pages('<p><strong>Páginas:</strong> ', '</p>', 'número'); ?>

		<?php endwhile; endif; ?>

		<h4><center><a href="javascript:history.go(-1)">&laquo;&laquo; Voltar</a> | <a href="#">Topo</a> <?php edit_post_link('(editar)'); ?></center></h4>

	</div>
</p>

<?php comments_template(); ?>

<?php endwhile; else: ?>
<p><?php _e('Lamento, página não encontrada.'); ?></p>
<?php endif; ?>

<?php get_footer(); ?>

Line 22 is <?php endwhile; else: ?>

It was working fine until recently, and I haven’t touched these php files in ages, so I have no clue as to why this happened now. Does anyone have an idea of what’s wrong?
Thanks.