Brilliant! Works a treat, thanks Caroline!!
I also just noticed that the offending effect stops happening when the page is refreshed.
Here’s the url
I’ve removed the duplicates and, so you can see what I mean, the following line:
filter: url(inc/desaturate.svg#greyscale); /* Firefox */
To see the issue, brush the pointer over some of the circles when using Firefox and see if they disappear.
Firefox seems to be updated (v 18.0.2)